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Jillus.Because he is too Jillus (jealous) he couldn't sleep, then decides to apply "NO DREAMING" rule.Also, he has a hard head.
My opinion based on how I fought them in game.Ghadius:Big targetPredictable movesHas 1 form onlyKing of Sorrow:Moving target in the 1st battle but in the 2nd battle you need to hit the weak points of his barrier, which is also moving btwUnpredictable movesHas 2 formsSo, King of Sorrow is the best villain for me and he's more of a challenge than Ghadius. That is all
Ghadius isn't even the final boss of DTP though. Nahatomb is. With him you have to constantly keep moving to avoid his armored moos (if you don't currently have a moo in your hand) and his tongue and make sure you load all six canons. He has THREE stages, a first canon stage, a stage where you fight his insides after he eats you, and a second canon stage after he spits you up. If I remember correctly (I probably don't, I don't replay boss levels all too much I'm more interested in the levels themselves) you have to do all three stages without dying?? But even if you don't each stage is still decently challenging. There was a hell of a lot more multitasking in DTP than there was LV.